Correct option is A
The correct substitution of the highlighted words is
(a) should speak.
Explanation: The original sentence lacks proper structure. "He who has suffered most in the cause" is a conditional clause and must be followed by an appropriate suggestion like
should speak.
Grammatical rule used:
�� When giving a suggestion or recommendation, use "should" + base verb.
Example: Anyone who disagrees should speak now.
��
Information Booster: Avoid imperatives in complex conditional structures.